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Table of contents
1 Synthesis
1.1 Graphical modeling of metal bioavailability
1.2 How to assess soil metal availability?
1.3 Biomarkers as indicators of metal bioavailability to earthworm?
1.4 Environmental bioavailability, a key step in the causal model
1.5 Concluding remarks
2 Experimental design
2.1 Specication and assumptions of the structural equation model
2.2 Choice of the target earthworm species
2.3 Selection of the series of eld-contaminated soils
2.4 Choice of experimental conditions of the earthworm exposure
3 Subcellular partitioning of metals in earthworm and metal speciation in soils
3.1 Introduction
3.2 Material and Methods
3.3 Results and Discussion
3.4 Conclusions
4 Relationship between earthworm energy reserves and soil metal availability
4.1 Introduction
4.2 Materials and Methods
4.3 Results
4.4 Discussion
4.5 Conclusion
5 Internal metal compartmentalization and biomarkers in earthworms
5.1 Introduction
5.2 Methods
5.3 Results and Discussion
6 Testing the structural equation model of metal bioavailability to earthworm
6.1 Introduction
6.2 Model description
6.3 Methods
6.4 Results and discussion
6.5 Conclusions
Discussion & Perspectives
Bibliography

