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Table of contents
Chapter 1: General introduction
1.1 Foraging and breeding: the essentials of life
1.2 Seabird foraging
1.3 Timing of breeding in seabirds
1.4 New Zealand, a hotspot for winter-breeding seabirds
1.5 Research objectives and thesis structure
Chapter 2: Foraging ecology of a winter breeder, the Fiordland penguin
2.1 Abstract
2.2 Introduction
2.3 Materials and methods
2.3.1 Study site and field procedures
2.3.2 Data handling and processing
2.3.3 Statistical analyses
2.4 Results
2.4.1 At-sea movements and habitat use
2.4.2 Diving behaviour and foraging effort
2.4.3 Trophic level and isotopic niche width
2.5 Discussion
2.5.1 At-sea movements and habitat use
2.5.2 Diving behaviour and foraging effort
2.5.3 Trophic level and isotopic niche width
2.6 Supplementary material
Chapter 3: Fine scale foraging behaviour of southern Buller’s albatross, the only Thalassarche provisioning chicks through winter
3.1 Abstract
3.2 Introduction
3.3 Materials and methods
3.3.1 Study site and field procedures
3.3.2 Data handling and processing
3.3.3 Statistical analyses
3.4 Results
3.4.1 At-sea movements and activity budget
3.4.2 Foraging behaviour and trophic niche
3.5 Discussion
3.5.1 At-sea movements and activity budget
3.5.2 Foraging behaviour and trophic niche
3.6 Supplementary material
Chapter 4: Foraging niche overlap during chick-rearing in the sexually dimorphic Westland petrel
4.1 Abstract
4.2 Introduction
4.3 Materials and methods
4.3.1 Study site and field procedures
4.3.2 Data processing and statistical analyses
4.4 Results
4.4.1 At-sea movements and activity budgets
4.4.2 Fine-scale foraging behaviour
4.5 Discussion
4.5.1 At-sea movements and activity budget
4.5.2 Foraging behaviour and trophic niche
4.6 Supplementary material
Chapter 5: Niche segregation in New Zealand winter breeding seabirds
5.1 Abstract
5.2 Introduction
5.3 Materials and methods
5.3.1 Study sites and field procedures
5.3.2 Data processing and statistical analyses
5.4 Results
5.4.1 At-sea movements
5.4.2 Temporal pattern of foraging and diving behaviour
5.4.3 Diet and isotopic niche
5.5 Discussion
Chapter 6: General Discussion
6.1 Foraging patterns of winter- and summer- breeding species
6.2 Marine trophic resource in winter around New Zealand
6.3 Relation between winter-foraging and winter-breeding
6.4 Perspectives and future research


